Beloved Galápagos tortoise Gramma, the oldest resident of the San Diego Zoo, has died at about 141, the zoo says, adding that ...

Gramma, who died at the zoo on Thursday, was believed to have been born in 1884, when Chester A. Arthur was in the White ...

No one knows exactly when Gramma was born. But if the estimated birth year of 1884 is accurate, Chester Arthur occupied the ...

An occasional series where Mongabay founder Rhett Ayers Butler shares analysis, perspectives and story summaries. She moved ...

Gramma, a Galápagos tortoise described by her care specialists as “the queen of the zoo,” was euthanized last Thursday after ...

The San Diego Zoo's oldest resident, a Galapagos tortoise named Gramma, died Nov. 20. She was estimated to be 141 years old.

One Galápagos tortoise lived 141 years, turning quiet meals and slow steps into a legacy of care, memory, and conservation ...

Gramma the tortoise, the oldest living resident at the San Diego Zoo, died at the age of 141, according to the zoo.

After more than a century of munching on her favorite foods of romaine lettuce and cactus fruit, beloved Galapagos tortoise ...

The beloved Galapagos tortoise named Gramma that was the oldest resident of the San Diego Zoo at an estimated 141 years has ...

The San Diego Zoo announced Gramma, the Galápagos tortoise, died on Nov. 20. She was about 141 years old based on estimates ...

Gramma the tortoise, believed to be 141 years old and the oldest resident of the San Diego Zoo, was euthanized after suffering from progressive bone deterioration because of her age.